Ballenato hocicudo de Blainville - Mesoplodon densirostris (Blainville, 1817)

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Aguilar de Soto, N., Madsen, P. T., Tyack, P., Arranz, P., Marrero, J., Fais, A., Revelli, E., Johnson, M. (2012). No shallow talk: Cryptic strategy in the vocal communication of Blainville's beaked whales. Marine Mammal Science, 28 (2): E75-E92.

Allen, A. N., Schanze, J. J., Solow, A. R., Tyack, P. L. (2014). Analysis of a Blainville's beaked whale's movement response to playback of killer whale vocalizations. Marine Mammal Science, 30 (1): 154-168.  

Allen, B, M., Mead, J. G., Brownell, R. L. (2011). Species review of Blainville’s beaked whale, Mesoplodon densirostris. SC/63/SM16. 18 pp.

Arranz, P., Aguilar de Soto, N., Madsen, P. T., Brito, A., Bordes, F., Johnson, M. P. (2011). Following a foraging fish-finder: diel habitat use of Blainville’s beaked whales revealed by echolocation. Plos One, 6 (12):e28353.

Baird, R. W., Webster, D. L., McSweeney, D. J., Ligon, A. D., Schorr, G. S., Barlow, J. (2006). Diving behaviour of Cuvier's (Ziphius cavirostris) and Blainville's (Mesoplodon densirostris) beaked whales in Hawai‘i.Canadian Journal of Zoology, 84: 1120-1128.

Baird, R. W., Webster, D. L., Schorr, G. S., McSweeney, D. J., Barlow, J. (2008). Diel variation in beaked whale diving behaviour. Marine Mammal Science, 24 (3): 630-642.    

Bellido, J. J., Cabot, J., Farfán, M. A., Castillo, J. J., Martín, J. J., Mons, J. L., Muñoz, J. L., Vázquez, C., Real, R. (2008). New records of beaked whales (genus Mesoplodon) in south Spain. JMBA2-Biodiversity Records,

Besharse, J. C. (1971). Maturity and sexual dimorphism in skull, mandible, and teeth of beaked whale, Mesoplodon densirostris. Journal of Mammalogy, 52 (2): 297-315.    

Carrillo, M. (2003). Presence and distribution of the Ziphiidae family in the south west coast of Tenerife. Canary islands. 17 Conference of the Europen Cetacean Society. Las Palmas de Gran Canaria.

Carrillo, M., López-Jurado, L. F. (1998). Structure and behaviour of a Blainville’s beaked whale (Mesoplodon densirostris) group in Tenerife (Canary Islands). World Mar. Mamm. Sci. Conf. 20–24 January, 1998, Monaco.

Casinos, A., Filella, S. (1981). Notes on Cetaceans of the Iberian coasts: 4. A specimen of Mesoplodon densirostris (Cetacea, Hyperoodontidae) stranded on the Spanish Mediterranean littoral. Saugetierkundliche Mitteilungen, 29 (4): 61-67.

Casinos, A., Filella, S. (1995). Mesoplodon densirostris (Blainville, 1817) – Blainville-Zweizahnwal. Pp. 575-582. En: Robineau, D., Duguy, R., Klima, M. (Eds.). Band 6: Meeressäuger. Teil I: Whale und Delphine – Cetacea. Teil I B: Ziphiidae, Kogiidae, Physeteridae, Balaenidae, Balaenopteridae. En: Niethammer, J., Krapp, F. (Eds.). Handbuch der Säugetiere Europas. Aula Verlag, Wiesbaden.

Colom-Llavina, M. M. (2005). Metazoan parasites of marine mammals from the Caribbean and the western coast of North America. M.Sc. Thesis. University of Puerto Rico.

Covelo, P., Martínez, J. (2001). Varamientos de mamíferos marinos en las costas de España y Portugal entre 1996 y 1998: Atlancetus. Galemys, 13: 93-106.

Covelo, P., Martínez-Cedeira, J. A., Llavona, A., Díaz, J., López, A. (2016). Strandings of Beaked Whales (Ziphiidae) in Galicia (NW Spain) between 1990 and 2013. Journal of the Marine Biological Association of the United Kingdom, 96 (4): 925-931.

Cox, T. M., Ragen, T. J., Read, A. J., Vos, E., Baird, R. W., Balcomb, K., Barlow, J., Caldwell, J., Cranford, T., Crum, L., D’Amico, A., D’Spain, G., Fernández, A., Finneran, J., Gentry, R., Gerth, W., Gulland, F., Hilde- brand, J., Houser, D., Hullar, T., Jepson, P. D., Ketten, D., MacLeod, C. D., Moore, S., Mountain, D., Palka, D., Ponganis, P., Rommel, S., Rowles, T., Taylor, B., Tyack, P., Wartzok, D., Gisiner, R., Mead, J., and Benner, L. (2006). Understanding the impacts of anthropogenic sound on beaked whales. J. Cetacean Res. Manage., 7: 177–187.

Faerber, M. M., Baird, R. W. (2010). Does a lack of observed beaked whale strandings in military exercise areas mean no impacts have occurred? A comparison of stranding and detection probabilities in the Canary and main Hawaiian Islands. Marine Mammal Science, 26 (3): 602-613.
Fernández, A. (2004). Pathological findings in stranded beaked whales during the naval military manoeuvers near the Canary Islands. ECS Newsletter, 42 (Special issue): 37-40.

Fernández, A., Arbelo, M., Martín, V. (2013). No mass strandings since sonar ban. Nature, 497 (7449): 317.

Henderson, E. E., Martin, S. W., Manzano-Roth, R., Matsuyama, B. M. (2016). Occurrence and Habitat Use of Foraging Blainville's Beaked Whales (Mesoplodon densirostris) on a US Navy Range in Hawaii. Aquatic Mammals, 42 (4): 549-562.

Heyning, J. E. (1984). Functional morphology envolved in intraspecific fighting of the beaked whale, Mesoplodon carlhubbsi. Canadian Journal of Zoology, 62: 1645-1654.

Heyning, J. E., Mead, J. G. (1996). Suction feeding in beaked whales: morphological and observational evidence. Nat. Hist. Mus. Los Angeles County Contributions in Science, 464: 1-12.

Jefferson, T. A., Leatherwood, S., Webber, M. A. (1993). FAO species identification guide. Marine Mammals of the World. FAO, Rome. 320 pp.

Johnson, M., Madsen, P. T., Zimmer, W. M. X., Aguilar de Soto, N., Tyack, P. L. (2004). Beaked whales echolocate on prey. Proceedings of the Royal Society Biological Sciences Series B, 271 (Suppl 6): S383-S386.

Johnson, M., Madsen, P. T., Zimmer, W. M. X., de Soto, N. A., Tyack, P. L. (2006). Foraging Blainville's beaked whales (Mesoplodon densirostris) produce distinct click types matched to different phases of echolocation. Journal of Experimental Biology, 209 (24): 5038-5050.    

Law, R. J., Allchin, C. R., Jones, B. R., Jepson, P. D., Baker, J. R., Spurrier, C. J. H. (1997). Metals and organochlorines in tissues of a Blainville's beaked whale (Mesoplodon densirostris) and a killer whale (Orcinus orca) stranded in the United Kingdom. Marine Pollution Bulletin, 34 (3): 208-212.    

Law, R. J., Bennett, M. E., Blake, S. J., Allchin, C. R., Jones, B. R., Spurrier, C. J. H. (2001). Metals and organochlorines in pelagic cetaceans stranded on the coasts of England and Wales. Marine Pollution Bulletin, 42 (6): 522-526.

MacLeod, C. D. (2002). Possible functions of the ultradense bone in the rostrum of Blainville’s beaked whale (Mesplodon densirostris). Canadian Journal of Zoology, 80: 178-184

MacLeod, C. D., Claridge, D. E. (1998). Scarring in a living population of dense beaked whales, Mesoplodon densirostris, off the island of Great Abaco, The Bahamas. P. 84. En: Abstracts of the World Marine Mammal Science Conference, Monaco, 20–24 January 1998.

MacLeod, C. D., Santos, M. B., Pierce, G. J. (2003). A review of data on diets of beaked whales: Evidence of niche separation and geographic segregation between three genera. J. Mar. Biol. Assoc. UK, 83: 651-5.

MacLeod, C. D., Zuur, A. F. (2005). Habitat utilisation by Blainville’s beaked whales off Great Abaco, Northern Bahamas, in relation to seabed topography. Marine Biology, 147: 1-11.

Madsen, P. T., Aguilar de Soto, N., Arranz, P., Johnson, M. (2013). Echolocation in Blainville's beaked whales (Mesoplodon densirostris). Journal of Comparative Physiology A- Neuroethology Sensory Neural and Behavioral Physiology, 199 (6): 451-469.

McCann, C. (1965). The mesorostral groove in Ziphiidae, with special reference to its closure by ossification in Mesoplodon-Cetacea. Rec. Dom. Mus., 5: 83–88.

McSweeney, D. J., Baird, R. W., Mahaffy, S. D. (2007). Site fidelity, associations, and movements of Cuvier's (Ziphius cavirostris) and Blainville's (Mesoplodon densirostris)  beaked whales off the island of Hawai'i. Marine Mammal Science, 23 (3): 666-687.    

Mead, J. G. (1989a). Beaked whales of the genus Mesoplodon. Pp. 349-430. En: Ridgway, S. H., Harrison, R. (Eds.). Handbook of marine mammals. Volume 4. Academic Press, San Diego.

Mead, J. G. (1989b). Survey of reproductive data for the beaked whales (Ziphiidae). Rep. int. Whal. Comm. Special Issue, 6: 91-96.

Moore, J. C. (1968). Relationships among the living genera of beaked whales with classifications, diagnoses and keys. Fieldiana: Zoology, 53: 209-298.

Perrin, W. F., Myrick, A. C. (1980). Age Determination of Toothed Whales and Sirenians. Rep. Int. Whal. Comm. Special Issue 3.

Rankin, S., Barlow, J. (2007). Sounds recorded in the presence of Blainville’s beaked whales, Mesoplodon densirostris, near Hawai’i. J. Acoust. Soc. Am., 122 (1): 42-45.

Reiner, F. (1979). Nota sobre um raro ziphoid, Mesoplodon densirostris Blainville 1817, nas costas de Portugal. Memorias do Museu do Mar Serie Zoologica, 1 (4): 1-12.

Ritter, F., Brederlau, B. (1999). Behavioural observations of dense beaked whales (Mesoplodon densirostris) off La Gomera, Canary Islands (1995–1997). Aquatic Mammals, 25 (2): 55-61.

Rommel, S. A., Costidis, A. M., Fernández, A., Jepson, P. D., Pabst, D. A., McLellan, W. A., Houser, D. S., Cranford, T. W., van Helden, A. L., Allen, D. M., Barros, N. B. (2006).  Elements of beaked whale anatomy and diving physiology and some hypothetical causes of sonar-related stranding. J. Cetacean Res. Manag., 7 (3): 189-209.

Ross, G. J. B. (1984). The smaller cetaceans of the southeast coast of southern Africa. Ann. Cape Province Mus. (Nat. Hist.), 15: 173-410.

Saliki, J. T., Cooper, E. J., Rotstein, D. S., Caseltine, S. L., Pabst, D. A., McLellan, W. A., Govett, P., Harms, C., Smolarek, K. A., Romero, C. H.  (2006). A novel gammaherpesvirus associated with genital lesions in a Blainville's beaked whale (Mesoplodon densirostris). Journal of Wildlife Diseases, 42 (1): 142-148.    

Santos, M. B., Martín, V., Arbelo, M., Fernández, A., Pierce, G. J. (2007). Insights into the diet of beaked whales from the atypical mass stranding in the Canary Islands in September 2002. Journal of the Marine Biological Association of the United Kingdom, 87 (1): 243-251.

Secchi, E. R., Zarzur, S. (1990). Plastic debris ingested by a Blainville's beaked whale, Mesoplodon densirostris, washed ashore in Brazil. Aquatic Mammals, 25: 21-24.

Stanton, T., Lavery, A. C., Johnson, M. P., Madsen, P. T., Tyack, P. L.  (2008). Classification of broadband echoes from prey of a foraging Blainville's beaked whale. Journal of the Acoustical Society of America, 123 (3): 1753-1762.    

Taylor, B. L., Baird, R., Barlow, J., Dawson, S. M., Ford, J., Mead, J. G., Notarbartolo di Sciara, G., Wade, P., Pitman, R. L. (2011). Mesoplodon densirostris. En: IUCN Red List of Threatened Species. Version 2011.1. <>.

Tyack, P. L., Johnson, M., Soto, N. A., Sturlese, A., Madsen, P. T. (2006). Extreme diving of beaked whales. Journal of Experimental Biology, 209 (21): 4238-4253.    

Tyack, P. L., Zimmer, W. M. X., Moretti, D., Southall, B.L., Claridge, D. E., Durban, J. W., Clark, C. W., D'Amico, A., DiMarzio, N., Jarvis, S., McCarthy, E., Morrissey, R., Ward, J., Boyd, I. (2011). Beaked Whales Respond to Simulated and Actual Navy Sonar. PLOS ONE, 6 (3): e17009.    

Valverde, J. A. (1997). Notes on a specimen of Blainville's beaked whale Mesoplodon densirostris (De Blainville, 1817) stranded on the coast of Doñana, Huelva, southern Spain. European Research on Cetaceans, 10: 184-189.



Ismael Miján Vilasánchez
Sociedade Galega de Historia Natural
Apdo 356, Ferrol 15401 A Coruña

Alberto Suárez-Esteban
Estación Biológica de Doñana (CSIC)
Avda. Américo Vespucio s/n, 41013  Sevilla

Fecha de publicación: 22-06-2011

Revisiones: 29-09-2017

Miján, I., Suárez-Esteban, A. (2017). Ballenato hocicudo de Blainville – Mesoplodon densirostris. En: Enciclopedia Virtual de los Vertebrados Españoles. Sanz, J. J., Barja, I. (Eds.). Museo Nacional de Ciencias Naturales, Madrid.